Beheregaray, L. B., Gibbs, J. P., Havill, N., Fritts, T. H., Powell, J. R., & Caccone, A. (2004). Giant tortoises are not so slow: Rapid diversification and biogeographic consensus in the Galápagos. National Academy of Sciences.

Beheregaray, Luciano B., James P. Gibbs, Nathan Havill, Thomas H. Fritts, Jeffrey R. Powell, and Adalgisa Caccone. Giant Tortoises Are Not so Slow: Rapid Diversification and Biogeographic Consensus in the Galápagos. National Academy of Sciences, 2004.

Beheregaray, Luciano B., et al. Giant Tortoises Are Not so Slow: Rapid Diversification and Biogeographic Consensus in the Galápagos. National Academy of Sciences, 2004.