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Kruuk, Loeske E. B., Tim H. Clutton-Brock, Jon Slate, Josephine M. Pemberton, Sue Brotherstone, και Fiona E. Guinness. Heritability of Fitness in a Wild Mammal Population. The National Academy of Sciences, 2000.

Kruuk, Loeske E. B., et al. Heritability of Fitness in a Wild Mammal Population. The National Academy of Sciences, 2000.