Carabidae Charcoal Dominant species Ecological Niche Model Eucalyptus Euscaphis japonica Forest History GARP Landsat Last Glacial Maximum MaxEnt Maxent Mediterranean area Moscow Region Paleoecology Permutation test Phylogenetic distance Phylogenetic relationships Pinus mugo/uncinata Pinus nigra Pinus sylvestris Quantitative distribution Quercus pubescens Relative groups SDMtoolbox amphibian aridization association group beta diversity biodiversity